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Celebs You Totally Forgot Guest-Starred on 'Grey's Anatomy'

Josh Radnor

Season: 15
The How I Met Your Mother actor played John, a possible love interest for Meredith. It was going well until he said he didn’t like kids.

Alyssa Milano

Season: 16
The actress and her Charmed co-star Holly Marie Combs played sisters who had to decide whether or not to take their other sister off of life support.

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